* * * * *
THE VARIATION
OF
ANIMALS AND PLANTS
UNDER DOMESTICATION.
BY CHARLES DARWIN, M.A., F.R.S., &c.
IN TWO VOLUMES.--VOL. I.
WITH ILLUSTRATIONS.
LONDON:
JOHN MURRAY, ALBEMARLE STREET.
1868.
_The right of Translation is reserved._
* * * * *
BY THE SAME AUTHOR.
* * * * *
ON THE ORIGIN OF SPECIES BY MEANS OF NATURAL SELECTION; or The PRESERVATION of FAVOURED RACES in the STRUGGLE for LIFE. Fourth Edition (_Eighth Thousand_), with Additions and Corrections. 1866. ... MURRAY.
A NATURALIST'S VOYAGE ROUND THE WORLD; or, A JOURNAL OF RESEARCHES into the NATURAL HISTORY and GEOLOGY of the COUNTRIES visited during the Voyage of H.M.S. Beagle, under the Command of Capt. FITZ-ROY, R.N. _Tenth Thousand_. ... MURRAY.
ON THE STRUCTURE AND DISTRIBUTION OF CORAL REEFS. ... SMITH, ELDER, & Co.
GEOLOGICAL OBSERVATIONS ON VOLCANIC ISLANDS. ... SMITH, ELDER, & Co.
GEOLOGICAL OBSERVATIONS ON SOUTH AMERICA. ... SMITH, ELDER, & Co.
A MONOGRAPH OF THE CIRRIPEDIA. With numerous Illustrations. 2 vols. 8vo. ... HARDWICKE.
ON THE VARIOUS CONTRIVANCES BY WHICH BRITISH AND FOREIGN ORCHIDS ARE FERTILISED BY INSECTS; and on the GOOD EFFECTS of CROSSING. With numerous Woodcuts. ... MURRAY.
ON THE MOVEMENTS and HABITS of CLIMBING PLANTS. With Woodcuts. ... WILLIAMS & NORGATE.
* * * * *
LONDON: PRINTED BY WILLIAM CLOWES AND SONS, STAMFORD STREET, AND CHARING CROSS.
* * * * *
{iii}
CONTENTS OF VOLUME I.
INTRODUCTION ... Page 1
CHAPTER I.
DOMESTIC DOGS AND CATS.
ANCIENT VARIETIES OF THE DOG--RESEMBLANCE OF DOMESTIC DOGS IN VARIOUS COUNTRIES TO NATIVE CANINE SPECIES--ANIMALS NOT ACQUAINTED WITH MAN AT FIRST FEARLESS--DOGS RESEMBLING WOLVES AND JACKALS--HABIT OF BARKING ACQUIRED AND LOST--FERAL DOGS--TAN-COLOURED EYE-SPOTS--PERIOD OF GESTATION--OFFENSIVE ODOUR--FERTILITY OF THE RACES WHEN CROSSED--DIFFERENCES IN THE SEVERAL RACES IN PART DUE TO DESCENT FROM DISTINCT SPECIES--DIFFERENCES IN THE SKULL AND TEETH--DIFFERENCES IN THE BODY, IN CONSTITUTION--FEW IMPORTANT DIFFERENCES HAVE BEEN FIXED BY SELECTION--DIRECT ACTION OF CLIMATE--WATER-DOGS WITH PALMATED FEET--HISTORY OF THE CHANGES WHICH CERTAIN ENGLISH RACES OF THE DOG HAVE GRADUALLY UNDERGONE THROUGH SELECTION--EXTINCTION OF THE LESS IMPROVED SUB-BREEDS.
Table of contents (by pages)
- 1: The Variation of Animals and Plants Under Domestic
- 2: Pigs belong to two distinct types
- 3: Osteological differences in the skull
- 4: Extremity of the furcula of fowls
- 5: 1 thevariation of animals and plantsunder domestication
- 6: Or unconsciously and unintentionally
- 7: May justly be called incipient species
- 8: By the extermination of the older intermediate forms
- 9: Now this hypothesis may be tested
- 10: Undergoing modification in the course of their descent
- 11: Built on the same type or pattern
- 12: In considering the theory of natural selection
- 13: With this most ancient variety a pariah like dog coexisted
- 14: The character of this animal during the Neolithic period
- 15: As do the savages of Australia those of the wild Dingo
- 16: Apparently the Canis cancrivorus
- 17: From the time of Gueldenstaedt to that of Ehrenberg
- 18: The feral dogs of La Plata have not become dumb
- 19: But true black and tan greyhounds are excessively rare
- 20: Because their periods of gestation are different
- 21: And so useful a group as the Canidae
- 22: 61 the molar teeth stand obliquely
- 23: A fourth cuneiform bone is developed
- 24: An additional pair of molar teeth in either jaw
- 25: As with the greyhound in coursing hares
- 26: Says that the greyhound within the last fifty years
- 27: Our pointers are certainly descended from a Spanish breed
- 28: Blyth 89 remarks on this passage
- 29: This 47 animal was left for only eight weeks at Mombas
- 30: And extreme variability in colouring
- 31: So that one sees appearing by monstrosity
- 32: Very unlike their Spanish progenitors
- 33: For even asses are sometimes dappled
- 34: Fallow dun Devonshire pony fig
- 35: And grey Kattywar horses being striped
- 36: The stripes are plainest when the colt is first foaled
- 37: And the remaining ten were zains
- 38: And spinal stripes in the horse
- 39: According to Nathusius Schweineschaedel
- 40: As Ruetimeyer apparently suspects
- 41: Nathusius makes a remarkable statement Schweineschaedel
- 42: 160 Nathusius states positively s
- 43: In the common domestic boar as 13
- 44: The tusks and bristles reappear with feral boars
- 45: Feral pigs generally revert to that of the wild boar
- 46: Independently of their important osteological differences
- 47: For I am informed by Professor Ruetimeyer
- 48: As zebus inhabit a distant and much hotter region
- 49: And probably belonged to the primigenius type
- 50: In other parts of the Falkland Islands
- 51: Andersson has described 205 the Damara
- 52: And a niata bull and cow invariably produce niata calves
- 53: Such as the semi monstrous niata cattle
- 54: Scarcely inherited a single point of the long horned breed
- 55: Arise from a crest on the frontal bone
- 56: On the excellent authority of Hermann von Nathusius
- 57: The fleeces recover their soft quality
- 58: Especially in the case of the Mauchamp race
- 59: And their absence to be characteristic of the genus Capra
- 60: And for the richnesse of the skin
- 61: Different from that of the Angora
- 62: And their offspring were either blacks or chinchillas
- 63: And are then called Himalayans
- 64: Under the name of Lepus magellanicus
- 65: Brought home two of these feral rabbits in spirits of wine
- 66: The two little Porto Santo rabbits
- 67: Skull of large Lop eared Rabbit
- 68: The whole meatus is directed more forwards
- 69: Measured across the zygomatic arch
- 70: Both had eight lumbar vertebrae
- 71: Ninth and tenth dorsal vertebrae
- 72: The acromion sends out a rectangular bar
- 73: And of four large but not fattened lop eared rabbits
- 74: The three Porto Santo feral rabbits Nos
- 75: The seven large lop cared rabbits
- 76: According to the length of the Skull
- 77: Hence I infer that in the Angora breed
- 78: Conjoined probably with the disuse of their muscles
- 79: As in the classification of natural species
- 80: As we shall see in the next chapter
- 81: Pouters stand remarkably upright
- 82: And with that round the eyes bare and likewise carunculated
- 83: With some little carunculated skin over the nostrils
- 84: The Scanderoon has a very short
- 85: And only slightly carunculated
- 86: From these several descriptions we see that with Runts
- 87: European Fantails Pfauen Taube
- 88: The beak of the Turbit is very short
- 89: Or three summersaults at a time
- 90: Esquilant possessed a blue Baldhead
- 91: I have kept two varieties of Laughers
- 92: For it is described by Bechstein
- 93: Many of the genera of the Columbidae
- 94: Bult's magnificent collection of Pouters
- 95: These marks are generally rusty red before the first moult
- 96: On comparing the skeletons of Columba livia
- 97: And the premaxillary bones are proportionally broader
- 98: Bussorah Pigeon
- 99: The extremities of the furcula
- 100: Whilst in two carriers and in a runt the tongue
- 101: Tegetmeier has seen two young birds in the same nest
- 102: I have counted eight scutellae on the hind toe of a runt
- 103: Fantail by standard to end of tail 1
- 104: In comparison with the rock pigeon
- 105: As fancy pigeons are generally kept in small aviaries
- 106: And the whole furcula was proportionally shorter
- 107: The length of the scapulae and furcula
- 108: The size and shape of the perforations in the sternum
- 109: Skirving repeatedly saw whole flocks settle on the water
- 110: But is closely allied to true Columba
- 111: When this form is domesticated chequered birds appear
- 112: Affinis from the cliffs of England
- 113: The common dovecot pigeon is not chequered
- 114: Are descended from distinct aboriginal stocks
- 115: The hypothesis that such species formerly existed
- 116: We can understand why they have not become feral
- 117: And this is not a universal rule with the Columbidae
- 118: Esquilant has seen a chequered Runt
- 119: In seven some white feathers appeared on the croup
- 120: This likewise is the case with white fantails
- 121: Livia or from several aboriginal species
- 122: Carriers and short faced tumblers
- 123: 349 Akber Khan possessed seventeen distinct kinds
- 124: The extremely slight improvement in Pouters
- 125: That the Turbit is the Cortbeck of Aldrovandi
- 126: They are briefly alluded to by Willughby
- 127: Persica et Turcica of this author comes the nearest
- 128: The Turbit apparently without its frill
- 129: The fancier in the time of Aldrovandi
- 130: And that other fanciers sacrifice everything for plumage
- 131: In the strains long kept distinct by different fanciers
- 132: This has occurred in the case of the pouter
- 133: And runt fanciers not caring about the beak or wattle
- 134: Nor has the extreme limit of variability in the beak
- 135: Several breeds are abnormal in character
- 136: Feathers closely adpressed to the body
- 137: 364 Sub breed g Guelderlands
- 138: With black skin and periosteum
- 139: A breed were utterly neglected
- 140: The Gallus bankiva apparently fulfils every requirement
- 141: The Malayan males generally had a red ear lappet
- 142: That wild specimens of the Gallus bankiva
- 143: Between Gallus bankiva and the Game fowl
- 144: Silver spangled Hamburgh fowls
- 145: One subsequently assumed pale orange coloured hackles
- 146: One of them assumed yellowish white hackles
- 147: Tegetmeier has 244 also remarked to me
- 148: For the domestic fowl is forbidden
- 149: In 1600 Aldrovandi describes seven or eight breeds of fowls
- 150: The Cochins laying buff coloured eggs
- 151: Gold and silver pencilled Hamburghs
- 152: And the males have not hackles
- 153: With the exception of the Dorkings
- 154: He well describes the peculiarities of this protuberance
- 155: But rumpless fowls often produce chickens with tails
- 156: The tarsi are often feathered
- 157: Bankiva as the standard of comparison
- 158: And lengthened in the Cochin skull
- 159: The two ascending branches of the premaxillary
- 160: And the ascending branches of the premaxillary
- 161: Which are generally anchylosed together
- 162: Being hardly enlarged in the Bankiva
- 163: Or the changed outline of the occipital foramen
- 164: Bankiva being called a hundred
- 165: The weight of the wild Bankiva
- 166: Gallus bankiva male 4
- 167: But likewise with tufted ducks
- 168: Abdominal sack largely developed
- 169: As with hook billed and penguin ducks
- 170: As the loquacity of the Call duck is highly serviceable
- 171: The Penguin duck is the most remarkable of all the breeds
- 172: And the descending haemal processes are much shortened
- 173: In comparison with the wild duck
- 174: As well as their slightly lessened length
- 175: The prominence of the crest of the sternum
- 176: The shaft divides into filaments which acquire barbules
- 177: Thornton's stock of common and pied peacocks
- 178: Furnished with an ample white topknot
- 179: If two topknotted birds are matched
- 180: The anal fins are sometimes double
- 181: With the exception of the Ligurian race or species
- 182: And races differing much in the cocoon
- 183: Of the many caterpillars thus reared
- 184: Presque toutes les races a trois mues
- 185: And certain black caterpillars
- 186: De Candolle as quite unknown in their aboriginal condition
- 187: De Candolle has shown that this probably has seldom occurred
- 188: A Fuegian on board the Beagle
- 189: The Polynesians are so frequently lost on the ocean
- 190: That none of these Cerealia exist
- 191: As Godron 547 has remarked
- 192: He sowed winter wheat in spring
- 193: Thus Colonel Le Couteur 556 says
- 194: The peculiar small eared and small grained wheat
- 195: Maize has varied in an extraordinary and conspicuous manner
- 196: When Metzger published his book
- 197: Such as the Portuguese Couve Tronchuda
- 198: Recrossed with purple broccoli
- 199: According to the view adopted by Godron and Metzger
- 200: In the Pois geant sans parchemin
- 201: Thin skinned pods of an extraordinary shape
- 202: With respect to the varieties not naturally intercrossing
- 203: Produced extremely dissimilar tubers
- 204: 618 whilst others are extremely variable
- 205: The roso bears strong hardy leaves
- 206: But that the shaddock Citrus decumana
- 207: Which grew close to lemons and citrons
- 208: Peach and Nectarine Amygdalus Persica
- 209: Through clingstones of poor quality
- 210: A nectarine tree grew close by
- 211: But at Carclew 666 the unique case occurred
- 212: Clingstone and freestone peaches
- 213: And generally in the Hemskirke
- 214: With a somewhat elongated stone
- 215: And borne on a stout very short footstalk
- 216: Acerba and praecox or paradisiaca
- 217: From the seed of the Ribston Pippin
- 218: Decaisne raised many seedlings from four distinct kinds
- 219: But the hybrid offspring from the Hautbois
- 220: But it frequently produces hermaphrodites
- 221: The branches are more or less downy or spinose
- 222: In 1844 London weighed 35 dwts
- 223: Six species are now recognised in the genus Cucurbita
- 224: Naudin in describing Cucurbita pepo p
- 225: Sageret 755 and Naudin found that the cucumber C
- 226: That De Candolle considered it as specifically distinct
- 227: Pendula have reproduced the parent form
- 228: There is only a single pistil
- 229: All relating to the Pelargonium
- 230: Centifolia produce an abundance of seed
- 231: Seedlings from this flower were semi double
- 232: Viola Altaica seems to be a distinct form
- 233: The nectary hardly varied at all
- 234: Hyacinth Hyacinthus orientalis
- 235: As occurs with differently coloured Hydras
- 236: Are hybrids from the peach and nectarine
- 237: The lower berries were well coloured black Frontignans
- 238: Variegata a perfect flower of A
- 239: And the red or coccineum varieties
- 240: Besides the old single red moss rose
- 241: Curtis 861 on the old Aimee Vibert Noisette
- 242: Taken from such abnormal fronds
- 243: As is well exemplified in the Dandy pelargonium
- 244: So that the whole tuber assumes a new character
- 245: The common and purple laburnum
- 246: Purpureus in my garden contained three
- 247: Caspary ascertained that only 2
- 248: Adami is not an ordinary hybrid
- 249: The stock sprouted and produced the bizzarria
- 250: Speciosissimus and phyllanthus
- 251: Variegation is much influenced
- 252: Some of these united tubers produced white
- 253: Which was neither pure Banksian nor pure Devoniensis
- 254: Laxton sent me various other crossed peas slightly
- 255: Turning now to the genus Matthiola
- 256: When fertilised with its own pollen
- 257: We may admit that in most cases the swelling of the ovarium
- 258: Chesnut mare bore a hybrid to a quagga
- 259: In illustration of the origin of Cytisus adami
- 260: Are eminently liable to bud variation
- 261: Produced by seed a closely allied nectarine
- 262: When budded or grafted on another
- 263: This dog has been called a Thibetan mastiff
- 264: Nott and Gliddon 'Types of Mankind
- 265: Blyth under the signature of Zoophilus
- 266: His account is fully confirmed by Rengger
- 267: Smith in 'Naturalist's Library
- 268: Jukes' 'Excursion in and about Newfoundland
- 269: Mackinnon on 'The Falkland Islands
- 270: Light dun and mouse dun horses dappled
- 271: 235 describes two wild horses from Mexico as roan
- 272: For the truly feral pigs of Jamaica
- 273: According to Ruetimeyer 'Zahmen
- 274: Published by order of the Royal Agricult
- 275: 221 'Journal of the Asiatic Soc
- 276: Clark gives a ludicrous proof of this fact
- 277: See Delamer on 'Pigeons and Rabbits
- 278: And who has largely bred pigeons
- 279: 55 states that it is absent in two species of Columba
- 280: Eaton's edition 1858 of Moore
- 281: Barbs may properly be called short beaked carriers
- 282: I have crossed turbits with barbs
- 283: Turtur vulgaris and the Ectopistes
- 284: Whose four grandparents were a white turbit
- 285: 366 'Ornamental and Domestic Poultry
- 286: These were probably feral birds
- 287: 'Ornamental and Domestic Poultry
- 288: See 'Naturgeschichte Deutschlands
- 289: Bankiva could have weighed so much
- 290: Crawfurd on the 'Relation of Domest
- 291: And Dixon in his 'Ornamental Poultry' p
- 292: 503 'Manuel de l'Educateur de Vers a Soie
- 293: The seeds of four genera of grasses
- 294: Triticoides becomes converted into true wheat
- 295: 559 'Pflanzen der Pfahlbauten
- 296: 598 'Die Pflanzen der Pfahlbauten
- 297: 'Teoria della Riproduzione Vegetale
- 298: 656 'Correspondence of Linnaeus
- 299: 'Catalogue of the Fruit in the Garden of Hort
- 300: ' by Deputation of the Caledonian Hort
- 301: Doubleday in 'Gardener's Chron
- 302: 758 Loudon's 'Arboretum et Fruticetum
- 303: 797 Loudon's 'Gardener's Magazine
- 304: Other cases are given by Braun
- 305: 850 See also Loudon's 'Arboretum
- 306: Than those on the ungrafted Aria
- 307: 922 Gaertner 'Bastarderzeugung
- 308: Read before the International Hort
